Department of Biology

  • Alternate Address: Department of Biology
  • Email:
  • Education: Ph.D. University of Utah, 1969; joined CSU in 1975
  • Professional Activities: Biotic and abiotic factors affecting both structure and function of grassland ecosystems. Editorial Board: Frontiers in Ecology and the Environment
  • Lifetime Special Recognition and Awards: Elected Fellow of The American Association for the Advancement of Science. ISI Highly cited researcher
  • Current Interests/Activities in Retirement: (1) Continuing research on grassland ecology with former students and colleagues. (2) Travel. (3) Golf